Dharmachakra mudra is considered a representation of Buddha's teachings about the cosmic order from the heart. The teaching sets into motion the wheel of dharma.
The cycle formed with the fingers represents the wheel of dharma. The left hand, which faces the heart, represents the inner world while the right represents the surrounding world. In metaphysical terms, the circle represents the union of method and wisdom. The jewels of Buddhism, which are Buddha, dharma and sangha, are represented by the three stretched fingers of the left hand.
The middle finger of the right hand denotes the listeners of Buddha's teachings. The ring finger represents the realizers of Buddha's teachings. The little finger refers to mahayana, "the great vehicle," which is one of the branches of Buddhism.
Practicing dharmachakra mudra calms the mind and elevates the state of the mind to a higher plane. The mudra also improves focus and helps to maintain positive attitude.
Dharmachakra mudra can be practiced with padmasana while in meditation to maximize its benefits.